/* ── Reset & Base ─────────────────────────────────────────────────────── */
*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }
html { scroll-behavior: smooth; }
body {
  font-family: "Inter", system-ui, -apple-system, sans-serif;
  font-size: 16px;
  line-height: 1.55;
  background: var(--bg);
  color: var(--ink);
  min-height: 100vh;
  transition: background 0.2s, color 0.2s;
}
a { color: inherit; }
img, video { max-width: 100%; display: block; }
ul, ol { list-style: none; }

/* ── Theme Tokens ────────────────────────────────────────────────────── */
:root {
  --bg:    #ffffff;
  --bg2:   #f5f7f9;
  --ink:   #0a0d12;
  --ink2:  #4a5159;
  --ink3:  #8a9099;
  --rule:  rgba(10,13,18,0.10);
  --ruleS: rgba(10,13,18,0.18);
  --red:   #b8112f;
  --px: 56px;
  --card-w: 300px;
  --card-img-h: 325px;
}
[data-theme="dark"] {
  --bg:    #0b0d10;
  --bg2:   #13161b;
  --ink:   #eaecef;
  --ink2:  #a8aeb6;
  --ink3:  #6b7077;
  --rule:  rgba(234,236,239,0.10);
  --ruleS: rgba(234,236,239,0.18);
  --red:   #e0394a;
}
@media (max-width: 768px) {
  :root { --px: 24px; --card-w: 240px; --card-img-h: 250px; }
}

/* ── Typography helpers ──────────────────────────────────────────────── */
.font-mono { font-family: "JetBrains Mono", ui-monospace, monospace; }
.label {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.14em;
  text-transform: uppercase;
  color: var(--ink3);
}
.label-red { color: var(--red); }

/* ── Preloader ── exact match to current arcarolab.org ───────────────── */
#preloader {
  display: flex;
  justify-content: center;
  align-items: center;
  position: fixed;
  top: 0; left: 0;
  width: 100%; height: 100%;
  background-color: white;
  z-index: 9999;
  transition: opacity 0.5s ease;
}
#preloader.fade-out { opacity: 0; pointer-events: none; }

.loading-image {
  position: absolute;
  width: 5vw;
  height: 5vw;
  border-radius: 50%;
  animation: preloader-rotate 0.5s linear infinite, preloader-flicker 0.25s step-start infinite;
  object-fit: cover;
}
#preloader-img2 {
  animation: preloader-rotate 0.5s linear infinite, preloader-flicker 0.25s step-start infinite;
  animation-delay: 0.125s;
}
@keyframes preloader-rotate {
  from { transform: rotate(0deg); }
  to   { transform: rotate(360deg); }
}
@keyframes preloader-flicker {
  0%, 100% { opacity: 1; }
  50%       { opacity: 0; }
}

/* ── Toolbar ─────────────────────────────────────────────────────────── */
#toolbar {
  position: sticky;
  top: 0;
  z-index: 100;
  background: var(--bg);
  border-bottom: 1px solid var(--rule);
  padding: 20px var(--px);
  display: flex;
  align-items: center;
  justify-content: space-between;
  transition: background 0.2s;
}
#toolbar a.logo { display: flex; align-items: center; text-decoration: none; }
#toolbar a.logo img {
  height: 49px;
  width: auto;
  display: block;
  transition: filter 0.2s;
}
[data-theme="dark"] #toolbar a.logo img { filter: invert(1); }

/* nav-right holds nav links + theme toggle together */
.nav-right {
  display: flex;
  align-items: center;
  gap: 36px;
}
.nav-links {
  display: flex;
  align-items: center;
  gap: 36px;
}
.nav-links a {
  font-size: 17px;
  font-weight: 500;
  text-decoration: none;
  color: var(--ink);
  padding-bottom: 4px;
  border-bottom: 2px solid transparent;
  transition: color 0.15s, border-color 0.15s;
  white-space: nowrap;
}
.nav-links a:hover { opacity: 0.75; }
.nav-links a.current {
  color: var(--red);
  border-bottom-color: var(--red);
}

.theme-toggle {
  width: 36px; height: 36px;
  border-radius: 18px;
  border: 1px solid var(--ruleS);
  background: transparent;
  color: var(--ink);
  cursor: pointer;
  display: flex; align-items: center; justify-content: center;
  padding: 0;
  flex-shrink: 0;
  transition: border-color 0.15s, color 0.15s;
}
.theme-toggle:hover { border-color: var(--ink3); }
.theme-toggle svg { display: block; }

/* Hamburger (mobile) */
.hamburger {
  display: none;
  flex-direction: column;
  gap: 5px;
  cursor: pointer;
  background: none;
  border: none;
  padding: 4px;
  color: var(--ink);
}
.hamburger span {
  display: block; width: 22px; height: 2px;
  background: var(--ink);
  transition: transform 0.2s, opacity 0.2s;
}

@media (max-width: 900px) {
  .nav-right { gap: 16px; }
  .nav-links {
    display: none;
    position: fixed;
    inset: 0;
    top: 106px;
    background: var(--bg);
    flex-direction: column;
    align-items: flex-start;
    gap: 0;
    padding: 24px var(--px);
    border-top: 1px solid var(--rule);
    z-index: 99;
  }
  .nav-links.open { display: flex; }
  .nav-links a {
    font-size: 22px;
    padding: 16px 0;
    width: 100%;
    border-bottom: 1px solid var(--rule);
    border-left: none;
  }
  .hamburger { display: flex; }
}

/* ── Footer ──────────────────────────────────────────────────────────── */
footer {
  padding: 28px var(--px) 32px;
  border-top: 1px solid var(--rule);
  display: flex;
  justify-content: space-between;
  align-items: center;
  font-size: 13px;
  color: var(--ink3);
  flex-wrap: wrap;
  gap: 8px;
}
footer .footer-left {
  display: flex;
  gap: 10px;
  align-items: center;
  flex-wrap: wrap;
}
footer .footer-name { color: var(--ink); font-weight: 600; }

/* ── Page sections ───────────────────────────────────────────────────── */
.px { padding-left: var(--px); padding-right: var(--px); }
.divider { border: none; border-top: 1px solid var(--rule); margin: 0 var(--px); }

/* ── HERO ────────────────────────────────────────────────────────────── */
.hero {
  position: relative;
  border-bottom: 1px solid var(--rule);
  overflow: hidden;
}
.hero-bg {
  position: absolute;
  top: 0; right: 0;
  width: 62%; height: 100%;
  object-fit: cover;
  object-position: center;
  display: block;
}
/* No dark-mode opacity reduction — image already has dark background */
.hero-fade-h {
  position: absolute; inset: 0;
  /* Left: fade from page-bg to transparent  Right: fade back to page-bg so black edge disappears */
  background: linear-gradient(90deg,
    var(--bg) 0%,
    var(--bg) 36%,
    rgba(255,255,255,0) 58%,
    rgba(255,255,255,0) 80%,
    var(--bg) 100%
  );
  transition: background 0.2s;
}
[data-theme="dark"] .hero-fade-h {
  background: linear-gradient(90deg,
    var(--bg) 0%,
    var(--bg) 36%,
    rgba(11,13,16,0) 58%,
    rgba(11,13,16,0) 80%,
    var(--bg) 100%
  );
}
.hero-fade-v {
  position: absolute; inset: 0;
  background: linear-gradient(180deg, rgba(255,255,255,0) 70%, var(--bg) 100%);
  transition: background 0.2s;
}
.hero-content {
  position: relative;
  padding: 96px var(--px) 88px;
  max-width: 920px;
}
.hero h1 {
  font-size: 40px;
  font-weight: 500;
  line-height: 1.15;
  letter-spacing: -0.8px;
  color: var(--ink);
  text-wrap: balance;
  max-width: 640px;
}
.hero-body {
  margin-top: 28px;
  font-size: 18px;
  line-height: 1.55;
  color: var(--ink2);
  text-wrap: pretty;
  max-width: 720px;
}
.hero-caption {
  margin-top: 28px;
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  color: var(--ink3);
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 500;
}
@media (max-width: 768px) {
  .hero-bg { width: 100%; height: 40%; top: auto; bottom: 0; opacity: 0.3; }
  .hero-fade-h { background: linear-gradient(180deg, var(--bg) 40%, rgba(255,255,255,0) 100%); }
  [data-theme="dark"] .hero-fade-h { background: linear-gradient(180deg, var(--bg) 40%, rgba(11,13,16,0) 100%); }
  .hero h1 { font-size: 32px; letter-spacing: -0.8px; }
  .hero-body { font-size: 16px; }
}

/* ── Carousel ────────────────────────────────────────────────────────── */
.carousel-section { padding: 80px 0; }
.carousel-header {
  display: flex;
  justify-content: space-between;
  align-items: flex-end;
  margin-bottom: 40px;
  padding: 0 var(--px);
}
.carousel-header h2 {
  font-size: 40px;
  font-weight: 500;
  letter-spacing: -1.1px;
  color: var(--ink);
}
.carousel-controls { display: flex; align-items: center; gap: 12px; }
.carousel-btn {
  width: 44px; height: 44px;
  border-radius: 22px;
  border: 1px solid var(--ruleS);
  background: var(--bg);
  color: var(--ink);
  font-size: 18px;
  cursor: pointer;
  display: flex; align-items: center; justify-content: center;
  transition: background 0.18s, color 0.18s, border-color 0.18s;
  user-select: none;
}
/* All buttons identical by default — next loses its inverted style */
.carousel-btn:hover { border-color: var(--ink3); }
/* Click flash: instant invert on press, fades back on release */
.carousel-btn:active,
.carousel-btn.btn-clicked {
  background: var(--ink) !important;
  color: var(--bg) !important;
  border-color: var(--ink) !important;
  transition: none !important;
}
.carousel-btn:disabled { opacity: 0.3; cursor: default; }
.carousel-link {
  margin-left: 14px;
  font-size: 14px;
  color: var(--ink2);
  font-weight: 500;
  text-decoration: none;
}
.carousel-link:hover { color: var(--ink); }

/* Scrollable viewport */
.carousel-viewport {
  overflow: hidden;
  padding: 0 var(--px);
}
.carousel-track {
  display: flex;
  gap: 28px;
  transition: transform 0.38s cubic-bezier(.4,0,.2,1);
  will-change: transform;
}

/* Each card — uniform width + fixed-height image */
.carousel-card { width: var(--card-w); flex: 0 0 auto; }
.carousel-card-img {
  width: var(--card-w);
  height: var(--card-img-h);
  overflow: hidden;
  border-radius: 4px;
  border: 1px solid var(--rule);
  background: var(--bg2);
  display: flex;
  align-items: center;
  justify-content: center;
}
[data-theme="dark"] .carousel-card-img { background: #e8eaed; }
.carousel-card-img img {
  width: 100%;
  height: 100%;
  object-fit: contain;   /* show full image, no cropping */
  display: block;
}
.carousel-card-img.no-img {
  color: var(--ink3);
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 500;
  text-align: center;
  padding: 16px;
}

.carousel-meta {
  margin-top: 16px;
  display: flex;
  align-items: baseline;
  gap: 12px;
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 600;
}
.carousel-year { color: var(--red); }
.carousel-journal { color: var(--ink3); white-space: nowrap; overflow: hidden; text-overflow: ellipsis; }
.carousel-title {
  margin-top: 8px;
  font-size: 16px;
  font-weight: 500;
  line-height: 1.3;
  color: var(--ink);
  text-wrap: pretty;
  display: -webkit-box;
  -webkit-line-clamp: 3;
  -webkit-box-orient: vertical;
  overflow: hidden;
}
.carousel-author {
  margin-top: 6px;
  font-size: 12px;
  color: var(--ink3);
  white-space: nowrap;
  overflow: hidden;
  text-overflow: ellipsis;
}
.carousel-track { cursor: grab; user-select: none; }
.carousel-track:active { cursor: grabbing; }
/* Linked paper title in carousel */
.carousel-title-link {
  color: inherit;
  text-decoration: underline;
  text-decoration-color: var(--rule);
  text-underline-offset: 3px;
  transition: text-decoration-color 0.15s;
}
.carousel-title-link:hover { text-decoration-color: var(--red); }
.dot { height: 3px; border-radius: 2px; background: var(--rule); transition: width 0.3s, background 0.3s; width: 20px; cursor: pointer; }
.dot.active { width: 48px; background: var(--red); }

@media (max-width: 768px) {
  .carousel-header { flex-direction: column; align-items: flex-start; gap: 16px; }
  .carousel-header h2 { font-size: 28px; }
}

/* ── Lab News + Conte ────────────────────────────────────────────────── */
.news-conte {
  padding: 72px var(--px) 64px;
  display: grid;
  grid-template-columns: 1.6fr 1fr;
  gap: 80px;
  align-items: start;
}
.news-conte h2 {
  font-size: 36px;
  font-weight: 500;
  letter-spacing: -1px;
  margin-bottom: 32px;
}

/* Lab News auto-scroll container */
.news-scroll {
  height: 480px;
  overflow: hidden;   /* hides scrollbar, JS animates scrollTop */
  border-top: 1px solid var(--rule);
  border-bottom: 1px solid var(--rule);
  position: relative;
}
/* inner content div — gets duplicated by JS for seamless loop */
#news-scroll-inner {
  /* no extra styles needed; JS drives scrollTop on parent */
}
.news-item {
  display: grid;
  grid-template-columns: 100px 1fr;
  gap: 24px;
  padding: 18px 0;
  border-bottom: 1px solid var(--rule);
}
.news-item:last-child { border-bottom: none; }
.news-item.transmission {
  display: block;
  padding: 14px 0;
  border-bottom: 1px solid var(--rule);
}
.news-date {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11.5px;
  color: var(--ink3);
  letter-spacing: 0.06em;
  padding-top: 2px;
  flex-shrink: 0;
}
.news-body { font-size: 15px; line-height: 1.55; color: var(--ink); text-wrap: pretty; }
.news-body a { color: var(--red); text-decoration: underline; text-underline-offset: 3px; }

/* Spacer after End of transmission lines — large gap before next loop's Please stand by */
.news-gap-spacer {
  height: 56px;
}
/* "Please stand by" item — no date column, full width, italic */
.news-item.news-standby {
  display: block;
  padding: 18px 0;
  border-bottom: 1px solid var(--rule);
  font-style: italic;
  color: var(--ink3);
}
/* End of transmission block */
.news-item.news-end-transmission {
  display: block;
  padding: 14px 0 20px;
  border-bottom: none;
}
.news-transmission-label {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 12px;
  font-weight: 600;
  letter-spacing: 0.14em;
  text-transform: uppercase;
  color: var(--ink3);
  margin-bottom: 12px;
}
.transmission-lines { margin-top: 6px; }
.transmission-lines .line {
  height: 2px;
  background: var(--ink3);
  margin: 5px 0;
}
.line-100 { width: 100%; }
.line-75  { width: 75%; }
.line-56  { width: 56.25%; }
.line-42  { width: 42.19%; }
.line-32  { width: 31.64%; }

.conte-card {
  padding: 28px;
  border: 1px solid var(--ruleS);
  border-radius: 4px;
  background: var(--bg2);
}
.conte-card .label { margin-bottom: 14px; }
.conte-title {
  font-size: 26px;
  font-weight: 600;
  line-height: 1.15;
  letter-spacing: -0.4px;
  margin-bottom: 14px;
}
.conte-body { font-size: 14.5px; line-height: 1.55; color: var(--ink2); margin-bottom: 20px; }
.conte-link {
  font-size: 14px;
  color: var(--red);
  font-weight: 500;
  text-decoration: none;
  border-bottom: 1px solid var(--red);
  padding-bottom: 2px;
}
.conte-link:hover { opacity: 0.8; }

@media (max-width: 900px) {
  .news-conte { grid-template-columns: 1fr; gap: 48px; }
}

/* ── Research Page ───────────────────────────────────────────────────── */
.research-header { padding: 96px var(--px) 72px; }
.research-header h1 {
  font-size: 62px;
  font-weight: 500;
  letter-spacing: -1.8px;
  line-height: 1.0;
  max-width: 1100px;
  text-wrap: balance;
}
.research-header p {
  margin-top: 36px;
  font-size: 18px;
  line-height: 1.55;
  color: var(--ink2);
  max-width: 820px;
  text-wrap: pretty;
}


/* Research page header background image */
body[data-page="research"] .research-header {
  position: relative;
  overflow: hidden;
  border-bottom: 1px solid var(--rule);
}
body[data-page="research"] .research-header::before {
  content: "";
  position: absolute;
  inset: 0;
  background-image:
    linear-gradient(90deg,
      var(--bg) 0%,
      var(--bg) 42%,
      rgba(255,255,255,0) 72%,
      var(--bg) 100%),
    linear-gradient(180deg,
      rgba(255,255,255,0) 70%,
      var(--bg) 100%),
    url('../img/research-img/macaque_fetal.png');
  background-repeat: no-repeat;
  background-size: 100% 100%, 100% 100%, auto 86%;
  background-position: center, center, right 7% center;
  opacity: 0.34;
  pointer-events: none;
}
[data-theme="dark"] body[data-page="research"] .research-header::before {
  background-image:
    linear-gradient(90deg,
      var(--bg) 0%,
      var(--bg) 42%,
      rgba(11,13,16,0) 72%,
      var(--bg) 100%),
    linear-gradient(180deg,
      rgba(11,13,16,0) 70%,
      var(--bg) 100%),
    url('../img/research-img/macaque_fetal.png');
  opacity: 0.28;
}
body[data-page="research"] .research-header > * {
  position: relative;
  z-index: 1;
}
@media (max-width: 768px) {
  body[data-page="research"] .research-header::before {
    background-size: 100% 100%, 100% 100%, auto 58%;
    background-position: center, center, right -8% bottom 8%;
    opacity: 0.18;
  }
}

.research-pills { margin-top: 36px; display: flex; gap: 12px; flex-wrap: wrap; }
.pill {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 12px;
  font-weight: 600;
  letter-spacing: 0.1em;
  text-transform: uppercase;
  padding: 8px 16px;
  border: 1px solid var(--ruleS);
  border-radius: 999px;
  text-decoration: none;
  color: var(--ink2);
  transition: border-color 0.15s, color 0.15s;
  white-space: nowrap;
}
.pill:hover { border-color: var(--ink3); color: var(--ink); }
.pill-red { border-color: var(--red); color: var(--red); }
.pill-red:hover { opacity: 0.8; }

.research-section {
  padding: 80px var(--px);
  border-top: 1px solid var(--rule);
}
.research-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 80px;
  align-items: start;
}
.research-section.reverse .research-grid { direction: rtl; }
.research-section.reverse .research-grid > * { direction: ltr; }

.research-section h2 {
  font-size: 56px;
  font-weight: 500;
  letter-spacing: -1.5px;
  line-height: 1.0;
}
.research-headline {
  margin-top: 24px;
  font-size: 22px;
  line-height: 1.35;
  font-weight: 400;
  font-style: italic;
  color: var(--ink2);
  letter-spacing: -0.2px;
  text-wrap: balance;
}
.research-body { margin-top: 20px; font-size: 17px; line-height: 1.6; color: var(--ink2); text-wrap: pretty; }
.research-pubs { margin-top: 36px; }
.research-pubs .label { margin-bottom: 14px; }
.pub-row {
  display: block;
  padding: 14px 0;
  border-top: 1px solid var(--rule);
  text-decoration: none;
  color: var(--ink);
}
.pub-row:hover { opacity: 0.75; }
.pub-row-meta {
  display: flex;
  align-items: baseline;
  gap: 12px;
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 600;
}
.pub-row-meta .year { color: var(--red); }
.pub-row-meta .journal { color: var(--ink3); }
.pub-row-title {
  margin-top: 4px;
  font-size: 16px;
  line-height: 1.4;
  font-weight: 500;
  text-wrap: pretty;
}
/* Linked title — matches publications page underline style */
.pub-row-title-link {
  color: inherit;
  text-decoration: underline;
  text-decoration-color: var(--rule);
  text-underline-offset: 3px;
  transition: text-decoration-color 0.15s;
}
.pub-row-title-link:hover {
  text-decoration-color: var(--red);
}
.pub-row-author { margin-top: 2px; font-size: 13px; color: var(--ink3); }

.research-media { width: 100%; }
.research-media-wrap {
  width: 100%;
  overflow: hidden;
  border-radius: 4px;
  border: 1px solid var(--rule);
  background: var(--bg2);
}
[data-theme="dark"] .research-media-wrap { background: #fff; }
.research-media-wrap.video-wrap { background: #000; }
[data-theme="dark"] .research-media-wrap.video-wrap { background: #000; }
.research-media-wrap img,
.research-media-wrap video { width: 100%; height: auto; display: block; }
.research-media-caption {
  margin-top: 14px;
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  color: var(--ink3);
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 500;
}

@media (max-width: 900px) {
  .research-header h1 { font-size: 42px; letter-spacing: -1px; }
  .research-grid { grid-template-columns: 1fr; gap: 48px; direction: ltr !important; }
  .research-section h2 { font-size: 38px; }
  .research-section.reverse .research-grid { direction: ltr; }
}
@media (max-width: 768px) {
  .research-header { padding: 60px var(--px) 48px; }
  .research-header h1 { font-size: 34px; }
  .research-section { padding: 56px var(--px); }
}

/* ── People Page ─────────────────────────────────────────────────────── */
.people-header { padding: 72px var(--px) 64px; }
.people-header h2 {
  font-size: 36px;
  font-weight: 500;
  letter-spacing: -1px;
  margin-bottom: 32px;
}
.people-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 36px 24px;
  align-items: start;
}
.person-card {
  width: 100%;
  text-align: center;
}
/* Circular photo — 208px (30% larger than original 160px), border-radius 50% */
.person-photo {
  width: 208px;
  height: 208px;
  overflow: hidden;
  border-radius: 50%;
  border: 1px solid var(--rule);
  background: var(--bg2);
  margin: 0 auto 16px;
  flex-shrink: 0;
}
[data-theme="dark"] .person-photo { background: #1a1d22; }
.person-photo img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  display: block;
}
.person-name {
  margin-top: 0;
  font-size: 16px;
  font-weight: 600;
  letter-spacing: -0.1px;
}
.person-role {
  margin-top: 3px;
  font-size: 13px;
  color: var(--ink2);
  line-height: 1.4;
}
.person-note { font-size: 12px; color: var(--ink3); margin-top: 2px; font-style: italic; }
.person-links { margin-top: 8px; display: flex; flex-wrap: wrap; gap: 5px; justify-content: center; }
.person-link {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 10px;
  letter-spacing: 0.1em;
  text-transform: uppercase;
  font-weight: 600;
  padding: 3px 8px;
  border: 1px solid var(--ruleS);
  border-radius: 999px;
  color: var(--ink2);
  text-decoration: none;
  transition: border-color 0.15s, color 0.15s;
}
.person-link:hover { border-color: var(--ink3); color: var(--ink); }

.alumni-collab {
  padding: 72px var(--px);
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 80px;
}
.alumni-collab h2 {
  font-size: 32px;
  font-weight: 500;
  letter-spacing: -0.8px;
  margin-bottom: 28px;
}
.list-row {
  display: grid;
  grid-template-columns: 1fr auto;
  gap: 16px;
  padding: 8px 0;
  border-top: 1px solid var(--rule);
  align-items: baseline;
}
.list-row a { font-size: 16px; font-weight: 500; text-decoration: none; color: var(--ink); }
.list-row a:hover { text-decoration: underline; text-underline-offset: 3px; }
.list-row span.name { font-size: 16px; font-weight: 500; color: var(--ink); }
.list-row span.detail { font-size: 13px; color: var(--ink3); font-style: italic; text-align: right; }

@media (max-width: 900px) {
  .people-grid { grid-template-columns: repeat(3, 1fr); }  /* overridden at 960px below */
  .alumni-collab { grid-template-columns: 1fr; gap: 48px; }
}

/* ── Publications Page ───────────────────────────────────────────────── */
.pub-page { padding: 40px var(--px) 64px; max-width: calc(1100px + var(--px) * 2); }
/* Tab bar — shared between publications.html and conferences.html */
.pub-tab-bar {
  display: flex;
  align-items: center;
  gap: 4px;
  margin-bottom: 48px;
  border-bottom: 1px solid var(--rule);
  padding-bottom: 0;
}
.pub-tab-btn {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 12px;
  font-weight: 600;
  letter-spacing: 0.1em;
  text-transform: uppercase;
  padding: 10px 20px;
  background: none;
  border: none;
  border-bottom: 2px solid transparent;
  margin-bottom: -1px;
  color: var(--ink2);
  cursor: pointer;
  transition: color 0.15s, border-color 0.15s;
}
.pub-tab-btn:hover { color: var(--ink); }
.pub-tab-btn.active {
  color: var(--red);
  border-bottom-color: var(--red);
}
.pub-scholar-link {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.1em;
  text-transform: uppercase;
  color: var(--ink3);
  text-decoration: none;
  margin-left: auto;
  padding: 10px 0;
  transition: color 0.15s;
}
.pub-scholar-link:hover { color: var(--ink); }

.pub-group { margin-bottom: 40px; display: grid; grid-template-columns: 160px 1fr; gap: 32px; align-items: start; }
.pub-year {
  font-size: 28px;
  font-weight: 500;
  letter-spacing: -0.6px;
  font-variant-numeric: tabular-nums;
  position: sticky;
  top: 110px;
}
.pub-list { border-top: 1px solid var(--rule); }
.pub-entry { padding: 14px 0; border-bottom: 1px solid var(--rule); font-size: 15.5px; line-height: 1.55; text-wrap: pretty; }
.pub-entry em { font-style: italic; color: var(--ink2); }
.pub-entry a {
  color: inherit;
  text-decoration: underline;
  text-decoration-color: var(--rule);
  text-underline-offset: 3px;
  transition: text-decoration-color 0.15s;
}
.pub-entry a:hover { text-decoration-color: var(--red); }

@media (max-width: 768px) {
  .pub-group { grid-template-columns: 1fr; gap: 16px; }
  .pub-year { font-size: 22px; position: static; }
  .pub-page { padding-top: 32px; }
}

/* ── Conferences Page ────────────────────────────────────────────────── */
.conf-entry { padding: 16px 0; border-bottom: 1px solid var(--rule); font-size: 15.5px; line-height: 1.55; text-wrap: pretty; color: var(--ink2); }
.conf-entry-author { color: var(--ink); }
.conf-entry-title { color: var(--ink); font-weight: 500; }

/* ── Open Science Page ───────────────────────────────────────────────── */
.project-card {
  padding: 64px var(--px);
  border-top: 1px solid var(--rule);
}
.project-card:first-child { border-top: none; }
.project-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 64px;
  align-items: start;
}
.project-card.reverse .project-grid { direction: rtl; }
.project-card.reverse .project-grid > * { direction: ltr; }

.project-card h2 {
  font-size: 48px;
  font-weight: 500;
  letter-spacing: -1.2px;
  line-height: 1.0;
}
.project-tagline {
  margin-top: 20px;
  font-size: 20px;
  line-height: 1.35;
  font-weight: 500;
  text-wrap: balance;
}
.project-desc {
  margin-top: 20px;
  font-size: 16px;
  line-height: 1.6;
  color: var(--ink2);
  text-wrap: pretty;
  max-width: 560px;
}
.project-links { margin-top: 18px; display: flex; flex-wrap: wrap; gap: 10px; }
.project-link {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 12px;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  font-weight: 600;
  padding: 8px 14px;
  border: 1px solid var(--ruleS);
  border-radius: 999px;
  color: var(--ink);
  text-decoration: none;
  transition: border-color 0.15s;
}
.project-link:hover { border-color: var(--ink3); }
.project-media {
  width: 100%;
  overflow: hidden;
  border-radius: 4px;
  border: 1px solid var(--rule);
  background: var(--bg);
}
[data-theme="dark"] .project-media { background: #fff; }
.project-media img { width: 100%; height: auto; display: block; }
.project-placeholder {
  width: 100%;
  aspect-ratio: 4 / 3;
  border: 1px dashed var(--rule);
  border-radius: 4px;
  display: flex;
  align-items: center;
  justify-content: center;
  color: var(--ink3);
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}

@media (max-width: 900px) {
  .project-grid { grid-template-columns: 1fr; direction: ltr !important; }
  .project-card.reverse .project-grid { direction: ltr; }
  .project-card h2 { font-size: 36px; }
}
@media (max-width: 768px) {
  .project-card { padding: 48px var(--px); }
}

/* end of stylesheet */

/* =============================================================
   RESPONSIVE OVERHAUL — additional breakpoints
   ============================================================= */

/* ── ~1100px: slightly tame large display text ── */
@media (max-width: 1100px) {
  .research-header h1 { font-size: 56px; letter-spacing: -1.4px; }
  .research-section h2 { font-size: 46px; }
  .project-card h2 { font-size: 42px; }
  .research-grid { gap: 56px; }
}

/* ── ~960px: pre-hamburger tightening (NO people grid change here) ── */
@media (max-width: 960px) {
  .hero h1 { font-size: 42px; letter-spacing: -1px; }
  .hero-content { padding: 72px var(--px) 64px; }
  .carousel-section { padding: 60px 0; }
  .research-header { padding: 72px var(--px) 56px; }
  .research-header h1 { font-size: 48px; }
  .research-section { padding: 60px var(--px); }
  .research-section h2 { font-size: 40px; }
  .research-grid { gap: 40px; }
  .people-header { padding: 56px var(--px) 48px; }
  /* People stays 3-col — 208px circles fit fine at this width */
  .alumni-collab { padding: 56px var(--px); gap: 48px; }
  .project-card { padding: 48px var(--px); }
  .project-grid { gap: 48px; }
  .news-conte { padding: 56px var(--px) 48px; gap: 56px; }
}

/* ── ~720px: people collapses to 2 cols (3 × 208px no longer fits) ── */
@media (max-width: 720px) {
  .people-grid { grid-template-columns: repeat(2, 1fr); gap: 28px 20px; }
  .person-photo { width: 160px; height: 160px; }
}

/* ── ~768px: mobile layout ── */
@media (max-width: 768px) {
  /* Hero */
  .hero h1 { font-size: 28px; letter-spacing: -0.6px; }
  .hero-content { padding: 56px var(--px) 48px; }
  .hero-body { font-size: 15px; }

  /* Carousel */
  .carousel-section { padding: 48px 0; }
  .carousel-header h2 { font-size: 26px; }
  .carousel-header { flex-direction: column; align-items: flex-start; gap: 14px; margin-bottom: 28px; }

  /* Research */
  .research-header { padding: 56px var(--px) 40px; }
  .research-header h1 { font-size: 32px; letter-spacing: -0.8px; }
  .research-section { padding: 48px var(--px); }
  .research-section h2 { font-size: 34px; }
  .research-headline { font-size: 18px; }
  .research-body { font-size: 15px; }

  /* People — already 2-col from 720px; shrink circles a touch more */
  .people-header { padding: 48px var(--px) 40px; }
  .person-photo { width: 140px; height: 140px; }
  .person-name { font-size: 15px; }

  /* Publications */
  .pub-group { grid-template-columns: 1fr; gap: 12px; }
  .pub-year { font-size: 20px; position: static; }

  /* Open Science */
  .project-card { padding: 40px var(--px); }
  .project-card h2 { font-size: 32px; letter-spacing: -0.8px; }
  .project-tagline { font-size: 17px; }

  /* News */
  .news-conte { padding: 48px var(--px) 40px; }
  .news-scroll { height: 360px; }
}

/* ── ~480px: very small phones ── */
@media (max-width: 480px) {
  .hero h1 { font-size: 24px; }
  .hero-content { padding: 40px var(--px) 36px; }

  /* News date: stack vertically so body text isn't squeezed */
  .news-item {
    grid-template-columns: 1fr;
    gap: 4px;
  }
  .news-date { padding-top: 0; }
  .news-scroll { height: 300px; }

  /* People: keep 2 cols but shrink circles */
  .person-photo { width: 110px; height: 110px; }
  .person-name { font-size: 13px; }
  .person-role { font-size: 11px; }

  /* Alumni/collab: tighter */
  .alumni-collab { padding: 40px var(--px); }
  .list-row { grid-template-columns: 1fr; gap: 2px; }
  .list-row span.detail { text-align: left; font-style: normal; }

  /* Carousel */
  .carousel-header h2 { font-size: 22px; }
  .carousel-controls { gap: 8px; }
  .carousel-link { margin-left: 6px; font-size: 12px; }
}

/* =============================================================
   EASTER EGG — Retro Mode (Konami Code)
   ============================================================= */

/* Retro pending — applied immediately on page load to prevent flash */
html.retro-pending body {
  background: #050a05 !important;
  color: #d4e8d4 !important;
}

/* ── Boot screen ── */
#retro-boot {
  position: fixed; inset: 0;
  background: #000;
  z-index: 99999;
  display: flex; align-items: center; justify-content: center;
  font-family: 'Press Start 2P', monospace;
  transition: opacity 0.4s ease;
}
.rb-inner { text-align: center; padding: 32px; max-width: 540px; width: 100%; }
.rb-stars { color: #555; font-size: 10px; letter-spacing: 6px; margin-bottom: 32px; }
.rb-t1 {
  font-size: 28px; color: #ffdd00;
  text-shadow: 3px 3px 0 #b89a00, 6px 6px 0 #6b5800;
  margin-bottom: 16px;
  animation: rb-flicker 4s infinite;
}
.rb-t2 { font-size: 10px; color: #00ff88; letter-spacing: 0.15em; margin-bottom: 10px; }
.rb-t3 { font-size: 7px; color: #555; letter-spacing: 0.1em; margin-bottom: 28px; }
.rb-divider { color: #1a1a1a; font-size: 10px; margin: 18px 0; letter-spacing: 1px; }
.rb-stats {
  font-size: 8px; color: #0af;
  margin: 20px 0 24px;
  display: flex; flex-direction: column; gap: 10px;
  text-align: left; display: inline-block;
}
.rb-stats > div { white-space: nowrap; }
.rb-stat-label { color: #ccc; }
.rb-dots { color: #333; }
.rb-stat-val { color: #ffdd00; }
.rb-press {
  font-size: 12px; color: #fff;
  animation: rb-blink 0.9s step-start infinite;
  margin: 28px 0 16px;
}
.rb-hint { font-size: 7px; color: #333; letter-spacing: 0.1em; }

@keyframes rb-blink { 0%,100%{opacity:1} 50%{opacity:0} }
@keyframes rb-flicker {
  0%,94%,100%{opacity:1} 95%{opacity:.7} 96%{opacity:1} 97%{opacity:.4} 98%{opacity:1}
}

/* ── CRT scanlines overlay ── */
body.retro-mode::after {
  content: '';
  position: fixed; inset: 0;
  background: repeating-linear-gradient(
    0deg, transparent 0, transparent 2px,
    rgba(0,0,0,0.13) 2px, rgba(0,0,0,0.13) 4px
  );
  pointer-events: none;
  z-index: 9997;
}

/* ── Global retro overrides ── */
/* Press Start 2P ≈ 1.3× visual weight of Inter at same px.
   Scale ≈ 0.78 keeps visual sizes close to the original layout. */
body.retro-mode {
  background: #050a05 !important;
  color: #d4e8d4 !important;
  font-family: 'Press Start 2P', monospace !important;
}
body.retro-mode * { font-family: 'Press Start 2P', monospace !important; }
body.retro-mode img { image-rendering: pixelated; }

/* ── Toolbar ── */
body.retro-mode #toolbar {
  background: #050a05 !important;
  border-bottom: 2px solid #00aa55 !important;
}
body.retro-mode .nav-links a { color: #d4e8d4 !important; font-size: 13px !important; letter-spacing: 0 !important; }
body.retro-mode .nav-links a.current { color: #ffdd00 !important; border-bottom-color: #ffdd00 !important; }
body.retro-mode .theme-toggle { border-color: #00aa55 !important; color: #00aa55 !important; }

/* ── Hero ── */
body.retro-mode .hero { background: #050a05 !important; }
body.retro-mode .hero-fade-h, body.retro-mode .hero-fade-v { opacity: 0.85 !important; }
body.retro-mode .hero-bg { filter: hue-rotate(90deg) saturate(2) brightness(0.25) !important; }
body.retro-mode .hero h1 {
  font-size: 16px !important; line-height: 2 !important;
  color: #ffdd00 !important; text-shadow: 2px 2px 0 #8a7000 !important; letter-spacing: 0 !important;
  max-width: none !important;
}
body.retro-mode .hero-body { font-size: 10px !important; line-height: 2.4 !important; color: #7ab87a !important; }
body.retro-mode .hero-caption { font-size: 8px !important; color: #3a5a3a !important; }

/* ── Carousel ── */
body.retro-mode .carousel-section { background: #050a05 !important; }
body.retro-mode .carousel-header h2 { font-size: 17px !important; color: #00ff88 !important; }
body.retro-mode .carousel-card-img { border: 3px solid #00aa55 !important; background: #000 !important; }
body.retro-mode .carousel-meta { font-size: 8px !important; }
body.retro-mode .carousel-year { color: #ffdd00 !important; }
body.retro-mode .carousel-journal { color: #0af !important; }
body.retro-mode .carousel-title { color: #d4e8d4 !important; font-size: 11px !important; line-height: 1.8 !important; }
body.retro-mode .carousel-author { color: #3a5a3a !important; font-size: 9px !important; }
body.retro-mode .dot { background: #1a2a1a !important; }
body.retro-mode .dot.active { background: #ffdd00 !important; }
body.retro-mode .carousel-btn { border-color: #00aa55 !important; color: #00aa55 !important; background: #050a05 !important; }
body.retro-mode .carousel-btn:active,
body.retro-mode .carousel-btn.btn-clicked { background: #00aa55 !important; color: #000 !important; border-color: #00aa55 !important; }
body.retro-mode .carousel-link { font-size: 9px !important; color: #7ab87a !important; }

/* ── News ── */
body.retro-mode .news-conte h2 { color: #00ff88 !important; font-size: 16px !important; }
body.retro-mode .news-scroll { border-color: #00aa55 !important; background: #000d00 !important; }
body.retro-mode .news-item { border-color: #0d1a0d !important; }
body.retro-mode .news-date { color: #00aa55 !important; font-size: 9px !important; }
body.retro-mode .news-body { color: #b8d4b8 !important; font-size: 10px !important; line-height: 2.2 !important; }
body.retro-mode .news-body a { color: #ffdd00 !important; }
body.retro-mode .news-transmission-label { color: #0af !important; font-size: 9px !important; }
body.retro-mode .transmission-lines .line { background: #0af !important; }
body.retro-mode .news-standby { color: #0af !important; font-style: normal !important; font-size: 10px !important; }

/* ── Conte card ── */
body.retro-mode .conte-card { background: #000d00 !important; border: 2px solid #ffdd00 !important; }
body.retro-mode .conte-title { color: #ffdd00 !important; font-size: 14px !important; }
body.retro-mode .conte-body { color: #7ab87a !important; font-size: 10px !important; line-height: 2.2 !important; }
body.retro-mode .conte-link { color: #00ff88 !important; border-color: #00ff88 !important; font-size: 9px !important; }
body.retro-mode .label { color: #0af !important; font-size: 8px !important; }

/* ── Research page ── */
body.retro-mode .research-header h1 { font-size: 22px !important; line-height: 1.8 !important; letter-spacing: 0 !important; color: #ffdd00 !important; }
body.retro-mode .research-header p  { font-size: 10px !important; line-height: 2.4 !important; color: #7ab87a !important; }
body.retro-mode .research-pills .pill { font-size: 8px !important; }
body.retro-mode .research-section h2 { font-size: 18px !important; color: #00ff88 !important; letter-spacing: 0 !important; }
body.retro-mode .research-headline { font-size: 12px !important; line-height: 2 !important; color: #7ab87a !important; font-style: normal !important; }
body.retro-mode .research-body { font-size: 10px !important; line-height: 2.4 !important; color: #7ab87a !important; }
body.retro-mode .research-pubs .label { font-size: 8px !important; color: #0af !important; }
body.retro-mode .pub-row-meta { font-size: 8px !important; }
body.retro-mode .pub-row-meta .year { color: #ffdd00 !important; }
body.retro-mode .pub-row-meta .journal { color: #0af !important; }
body.retro-mode .pub-row-title { font-size: 10px !important; line-height: 1.8 !important; }
body.retro-mode .pub-row-title-link { color: #d4e8d4 !important; text-decoration-color: #3a5a3a !important; }
body.retro-mode .pub-row-author { font-size: 8px !important; color: #3a5a3a !important; }
body.retro-mode .research-media-caption { font-size: 7px !important; color: #3a5a3a !important; }

/* ── People page ── */
body.retro-mode .people-header h2 { font-size: 16px !important; color: #00ff88 !important; }
body.retro-mode .person-name { font-size: 11px !important; color: #d4e8d4 !important; }
body.retro-mode .person-role { font-size: 9px !important; color: #7ab87a !important; }
body.retro-mode .person-note { font-size: 8px !important; color: #3a5a3a !important; }
body.retro-mode .person-link { font-size: 7px !important; border-color: #00aa55 !important; color: #7ab87a !important; }
body.retro-mode .alumni-collab h2 { font-size: 15px !important; color: #00ff88 !important; }
body.retro-mode .list-row a,
body.retro-mode .list-row span.name { font-size: 10px !important; color: #d4e8d4 !important; }
body.retro-mode .list-row span.detail { font-size: 8px !important; color: #3a5a3a !important; }

/* ── Publications / Conferences page ── */
body.retro-mode .pub-tab-btn { font-size: 9px !important; letter-spacing: 0 !important; }
body.retro-mode .pub-tab-btn.active { color: #ffdd00 !important; border-bottom-color: #ffdd00 !important; }
body.retro-mode .pub-scholar-link { font-size: 8px !important; color: #3a5a3a !important; }
body.retro-mode .pub-year { font-size: 17px !important; color: #00ff88 !important; }
body.retro-mode .pub-entry { font-size: 9px !important; line-height: 2.2 !important; color: #b8d4b8 !important; border-color: #0d1a0d !important; }
body.retro-mode .pub-entry a { color: #d4e8d4 !important; text-decoration-color: #3a5a3a !important; }
body.retro-mode .pub-entry em { color: #0af !important; }
body.retro-mode .conf-entry { font-size: 9px !important; line-height: 2.2 !important; color: #7ab87a !important; border-color: #0d1a0d !important; }
body.retro-mode .conf-entry-author { color: #d4e8d4 !important; }
body.retro-mode .conf-entry-title { color: #d4e8d4 !important; }

/* ── Tools & Data page ── */
body.retro-mode .project-card h2 { font-size: 18px !important; color: #ffdd00 !important; letter-spacing: 0 !important; }
body.retro-mode .project-tagline { font-size: 11px !important; color: #00ff88 !important; }
body.retro-mode .project-desc { font-size: 10px !important; line-height: 2.3 !important; color: #7ab87a !important; }
body.retro-mode .project-link { font-size: 8px !important; border-color: #00aa55 !important; color: #7ab87a !important; letter-spacing: 0 !important; }

/* ── Footer ── */
body.retro-mode footer { background: #050a05 !important; border-color: #00aa55 !important; }
body.retro-mode footer * { color: #3a5a3a !important; font-size: 9px !important; }

/* ── Dividers ── */
body.retro-mode .divider, body.retro-mode hr { border-color: #0d1a0d !important; }

/* ── 8-bit image effects in retro mode ── */

/* All images: nearest-neighbour scaling */
body.retro-mode img {
  image-rendering: pixelated !important;
  image-rendering: crisp-edges !important;
}

/* Pixel grid — 8px pitch matches PIX_FACTOR=8.
   Grid lines align perfectly with pixel block boundaries from the JS transform.
   Draws visible "seams" between each game pixel. */
body.retro-mode .carousel-card-img,
body.retro-mode .person-photo,
body.retro-mode .research-media-wrap,
body.retro-mode .project-media {
  position: relative;
  overflow: hidden;
}
body.retro-mode .carousel-card-img::after,
body.retro-mode .person-photo::after,
body.retro-mode .research-media-wrap::after,
body.retro-mode .project-media::after {
  content: '';
  position: absolute;
  inset: 0;
  background-image:
    repeating-linear-gradient(0deg,  rgba(0,0,0,0.22) 0, rgba(0,0,0,0.22) 1px, transparent 1px, transparent 8px),
    repeating-linear-gradient(90deg, rgba(0,0,0,0.22) 0, rgba(0,0,0,0.22) 1px, transparent 1px, transparent 8px);
  pointer-events: none;
  z-index: 3;
}

/* Colour filter: boost saturation + contrast to push hues toward
   a visually "limited palette" feel */
body.retro-mode .carousel-card-img img,
body.retro-mode .person-photo img,
body.retro-mode .research-media-wrap img,
body.retro-mode .project-media img {
  filter: saturate(2.8) contrast(1.5) brightness(0.85) !important;
}


/* ── Species shared element styles ──────────────────────────────── */
.species-label {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 11px; font-weight: 600; letter-spacing: 0.16em;
  text-transform: uppercase; color: var(--red); margin-bottom: 12px;
}
.species-name {
  font-size: 52px; font-weight: 500; letter-spacing: -1.3px; line-height: 1.0;
}
.species-latin {
  font-size: 18px; font-style: italic; color: var(--ink2);
  margin-top: 8px; letter-spacing: -0.2px;
}
.species-why {
  margin-top: 24px; font-size: 17px; line-height: 1.65;
  color: var(--ink2); text-wrap: pretty;
}

/* Retro overrides for species elements */
body.retro-mode .species-label { color: #0af !important; font-size: 8px !important; }
body.retro-mode .species-name { color: #ffdd00 !important; font-size: 18px !important; letter-spacing: 0 !important; }
body.retro-mode .species-latin { color: #00ff88 !important; font-size: 10px !important; font-style: normal !important; }
body.retro-mode .species-why { color: #7ab87a !important; font-size: 10px !important; line-height: 2.4 !important; }

/* ── Species comparison cards ────────────────────────────────────── */
.species-intro-p1 {
  font-size: 17px; line-height: 1.65; color: var(--ink);
  text-wrap: pretty; max-width: 860px;
}
.species-intro-p2 {
  font-size: 17px; line-height: 1.65; color: var(--ink2);
  text-wrap: pretty; max-width: 860px; margin-top: 24px;
}

.species-cards {
  padding: 72px var(--px) 64px;
  background: var(--bg2);
  border-top: 1px solid var(--rule);
  border-bottom: 1px solid var(--rule);
}
.species-card-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 48px;
  align-items: start;
}
.scard { display: flex; flex-direction: column; }
.scard-img-wrap {
  width: 140px; height: 140px;
  border-radius: 50%;
  overflow: hidden;
  border: 1px solid var(--rule);
  margin: 0 auto 20px;
  background: var(--bg);
  flex-shrink: 0;
}
.scard-img-wrap img { width: 100%; height: 100%; object-fit: cover; display: block; }
.scard-name {
  font-size: 20px; font-weight: 600; letter-spacing: -0.3px;
  text-align: center; color: var(--ink);
}
.scard-latin {
  font-size: 14px; font-style: italic; color: var(--ink3);
  text-align: center; margin-bottom: 20px;
}
.scard-section-label {
  font-family: "JetBrains Mono", ui-monospace, monospace;
  font-size: 10px; font-weight: 600; letter-spacing: 0.14em;
  text-transform: uppercase; color: var(--ink3);
  margin: 16px 0 8px;
}
.scard-list {
  list-style: none; padding: 0; margin: 0;
  display: flex; flex-direction: column; gap: 6px;
}
.scard-list li {
  font-size: 14px; line-height: 1.4; color: var(--ink);
  padding-left: 14px; position: relative;
}
.scard-list li::before {
  content: '↳';
  position: absolute; left: 0;
  color: var(--red); font-size: 11px;
}
.scard-list-constraints li { color: var(--ink2); }
.scard-list-constraints li::before { content: '↳'; color: var(--ink3); }

@media (max-width: 900px) {
  .species-card-grid { grid-template-columns: 1fr; gap: 48px; }
  .species-cards { padding: 56px var(--px); }
}

/* Retro overrides for cards */
body.retro-mode .species-cards { background: #000d00 !important; border-color: #00aa55 !important; }
body.retro-mode .scard-name { color: #ffdd00 !important; font-size: 11px !important; }
body.retro-mode .scard-latin { color: #00ff88 !important; font-size: 8px !important; font-style: normal !important; }
body.retro-mode .scard-section-label { color: #0af !important; font-size: 7px !important; }
body.retro-mode .scard-list li { color: #d4e8d4 !important; font-size: 8px !important; line-height: 2 !important; }
body.retro-mode .scard-list-constraints li { color: #7ab87a !important; }
body.retro-mode .scard-list li::before { color: #ffdd00 !important; }
body.retro-mode .scard-list-constraints li::before { color: #3a5a3a !important; }
body.retro-mode .species-intro-p1 { font-size: 10px !important; line-height: 2.4 !important; color: #d4e8d4 !important; }
body.retro-mode .species-intro-p2 { font-size: 10px !important; line-height: 2.4 !important; color: #7ab87a !important; }

/* ── Species detailed sections ───────────────────────────────────── */
.species-section {
  padding: 80px var(--px);
  border-top: 1px solid var(--rule);
}
.species-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 80px;
  align-items: start;
}
.species-grid > * { min-width: 0; } /* prevent long text/links blowing out grid cell */
.species-section.reverse .species-grid { direction: rtl; }
.species-section.reverse .species-grid > * { direction: ltr; }
.species-media { width: 100%; }
.species-media .species-img-wrap {
  width: 100%;
  height: auto;
  border-radius: 4px;
  border: 1px solid var(--rule);
  overflow: hidden;
  background: var(--bg2);
  margin: 0;
}
.species-media .species-img-wrap img {
  width: 100%; height: auto; object-fit: cover; display: block;
}

@media (max-width: 900px) {
  .species-grid { grid-template-columns: 1fr; gap: 40px; direction: ltr !important; }
  .species-section.reverse .species-grid { direction: ltr; }
  .species-section { padding: 56px var(--px); }
}

/* Retro overrides */
body.retro-mode .species-section { border-color: #0d1a0d !important; }
body.retro-mode .species-name { color: #ffdd00 !important; font-size: 18px !important; letter-spacing: 0 !important; }
body.retro-mode .species-latin { color: #00ff88 !important; font-size: 10px !important; font-style: normal !important; }
body.retro-mode .species-why { color: #7ab87a !important; font-size: 10px !important; line-height: 2.4 !important; }
body.retro-mode .species-label { color: #0af !important; font-size: 8px !important; }
body.retro-mode .species-media .species-img-wrap { border-color: #00aa55 !important; }

/* ── Species page title & phylogenetic tree ────────────────────── */
.species-page-title {
  font-size: 48px;
  font-weight: 500;
  letter-spacing: -1.2px;
  line-height: 1.05;
  margin-bottom: 32px;
  text-wrap: balance;
}

.species-intro-wrap {
  padding: 64px var(--px);
  display: grid;
  grid-template-columns: 1fr 460px;
  gap: 60px;
  align-items: start;
}
.species-intro-text {}
.species-tree-col { flex-shrink: 0; }
.species-tree-col svg { width: 100%; height: auto; display: block; }

@media (max-width: 1024px) {
  .species-intro-wrap { grid-template-columns: 1fr 340px; gap: 32px; }
}
@media (max-width: 800px) {
  .species-intro-wrap { grid-template-columns: 1fr; }
  .species-tree-col { max-width: 380px; }
}
@media (max-width: 960px) { .species-page-title { font-size: 36px; } }
@media (max-width: 768px) { .species-page-title { font-size: 28px; } }

/* Retro overrides */
body.retro-mode .species-page-title { font-size: 18px !important; color: #ffdd00 !important; letter-spacing: 0 !important; }